Source: Diamond Spheres: Hustle & Bustle

Incredibly durable and capable of travelling far without tiring, these coursers do not see nature as a place to thrive, but to endure.

Greater Expertise

The survivor has an improved talent progression, increasing from a Journeyman progression to a Virtuoso talent progression.

This alters skill expertise and replaces forage.

Hardened

At 1st level, the survivor gains the Body Control sphere and their choice of the Herbalism or Survivalism sphere as bonus skill talents.

This replaces survivalist.

Conditioned

At 3rd level, the survivor gains the benefits of the conditioning acclimation (as per the Navigation sphere (wilderness) package) as a constant effect, using their courser level as their associated ranks. This does not count towards the maximum number of acclimations the survivor can benefit from at once.

This replaces vigilant.

Rough Exterior

At 7th level, the survivor can always endure a failed Fortitude saving throw. Additionally, for as long as the survivor is meditating, they gain DR 1/— which stacks with similar sources, +1 per 5 courser levels they possess.

This replaces resourceful foraging.